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A friend of mine recently booked a multi-city one-way ticket covering three airlines that according to ITA Matrix includes this mysterious segment:

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Note that this is described as "First (F)", even though Jetstar is a low-cost carrier, doesn't offer more than Economy and Business, and their GDS guide doesn't recognize a fare class "F". The rest of the itinerary is also all "Economy (S)", which is a Starter Plus fare.

Any idea what kind of seat she's actually going to end up in, and how this happened?

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It isn't first class, it is a mistake made by ITA Software (unusually) in mapping the booking class (F) back to the cabin class (for JQ, booking class F is economy). A similar mistake is made by a few online travel agents for Aer Lingus which also uses F for economy.

Looking at ExpertFlyer it seems that JQ uses J and D for business; Y, B, V, T, S, R, Q, P, O, N, M, E, F, L, A, Z, I, K and H for economy, in order from most to least available.

The GDS guide you've linked is from 2014, they may have amended their codes since then; or it may be that the F class is not used for publicly sold fares.

You may want to report it to ITA as a bug.

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    Of all the letters in the alphabet to choose from, they just had to use F for a mid-cost economy fare didn't they? I mean, they knew Y and B were full-fare, so they weren't ignorant of the conventions used by everyone else in the industry; they just didn't care. – Zach Lipton Jan 13 '17 at 17:41

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